The epic story of Borderlands continues in part one of the new series, where fans find out what happens after Roland, Mordecai, Lilith, and Brick step out of Marcus's bus. As soon as they arrive in the once-prosperous town of Fyrestone, they realize that Pandora is a far more dangerous planet than they first imagined. The Vault Hunters will go up against a ruthless bandit known as "Nine Toes," but will they be able to overcome him?

The Vault Hunters have narrowly escaped Nine Toes and find themselves lost and unable to find the fabled “Vault.” Enter Patricia Tannis—a brilliant scientist who's struggling to cling to reality… and who is the Hunters' only hope of reaching the Vault. But can they trust her? Return to Pandora in this all-new, action-packed arc!

For the millions and millions of fans of the smash-hit video game BORDERLANDS, a lingering question exists: how did the Vault Hunters get on the bus with Marcus in the first place? Find out how Roland got on the bus with Mordecai, Lilith, and Brick in this exciting look at the soldiers of the Crimson Lance and what made him leave their ranks and search for the Vault. Surprises abound in the first part of a four-issue series chronicling where Borderlands began.